Oaks of the World

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  Quercus mannifera
Author Lindl. 1840 Bot. Reg. 26: t. 40
Synonyms

sessiliflora Salisb. var. cedrorum DC. 1864 pp
sessiliflora Salisb. var. pinnatifida Boiss. 1879

sessiliflora var. mannifera (Lindl.) Boiss. 1879
robur var. mannifera (Lindl.) Bruttini 1930
x mannifera O.Schwarz 1936
komarovii A.Camus 1939
atropatena O.Schwarz & Hess 1938
longifolia K.Koch 1849 nom. inval.
robur var. szovitzii A.DC 1864
robur var. araxina Trautv. 1873
araxina (Trautv)Grossh. 1930
x szovitzii (A.DC) Zabel 1903 illeg.

Local names
Range East Turquey to North Iran; 700-2000 m;
Growth habit shrub (2-6 m ); crown fastigiate; very branched;
Leaves 10-19 cm long, 4-9 cm wide; deciduous; thick; oval-oblong; both sides glabrous, dull grey-green above, yellowish green beneath; base truncate or slightly cordate, often asymmetrically; margin with 5-9 pairs of lobes with narrow sinuses; some stellate hairs near axils at the base; 7-8 vein pairs; few intercalary veins, near the foliar base; petiole 2.5-3.5 cm, hairless;
Flowers
Fruits cylindrical acorn, 2-3 cm long; peduncle absent or to 2 cm; flat cupule 2 cm in diameter, thick-walled; scales appressed, brown, subacute, oval, grey tomentose; maturing in 1 year;

Bark, twigs and
buds

grey bark remotely furrowed; twigs hairless; buds 5 mm long, glabrous; stipules deciduous, threadlike to lanceolate;
Hardiness zone, habitat hardy; good on silicious soils;
Miscellaneous -- N° 158 in A.Camus;
-- Sub-genus Quercus, Section Quercus, Series Roburoid;
-- The specific name is based on the presence on the leaves of a sugary substance, "manna", produced by action of aphids.
-- This taxon had often been, in the past, considered as a hybrid, mostly with Q. boissieri and Q. iberica ( or more exactly Q. infectoria subsp. veneris X Q. petraea subsp. polycarpa), but recent works (M. Avishai) enabled to clarify its taxonomical rank.

Subspecies and
varieties

2 varieties can be distinguished in nature:
-- var. mannifera
leaves oblong-elliptic with lobes truncate, shallow, narrow sinuses, without secondary lobes;

-- var. pinnatiloba (C.Koch) Avishai comb. and stat. nov.
leaves ovate, pinnatilobate, wide sinuses between subacute lobes, 1-3 secondary lobes at lobes margin;

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